Education

• B.S. 1999, Geography, Changchun College of Geology, Changchun, Jilin Province
• M.S. 2002, Quaternay Geology, Jilin University, Changchun, Jilin Province
• Ph.D. 2006, Quaternay Geology, Institute of Geology and Geophysics, Chinese Academy of Sciences, Beijing

Professional experiences

• 2009-present, Associate Research Professor, Institute of Geology and Geophysics, Chinese Academy of Sciences, Beijing
• 2006-2009, post-doc, Institute of Geology and Geophysics, Chinese Academy of Sciences, Beijing
• January-February 2008, November-December 2008, and November 2011, visiting scholar, Laboratoire de Meteorologie Dynamique, CNRS, Ecole Normale Superieure, Paris
• October 2025 and 2026, visiting scholar, University of Novi Sad, Serbia, to perform collaborative research of land snails in loess-paleosol sequences of southern Europe.

Academic Services and Honors

1. Head of the Mollusk Laboratory affiliated to the State Key Laboratory of Lithospheric Evolution and Environment
2. Member of the Life Evolution and Environment Committee, China Quaternary Research Association
3. Editorial Board Member, Scientific Reports
4. Editorial Board Member of ecology section, Biology
5. Peer Reviewer for Project Proposals, National Research, Development and Innovation Office (NRDIO), Hungary
6. Journal Reviewer for Communications Earth & Environment, GRL, Quaternary Science Reviews, Global and Planetary Change, Palaeogeography, Palaeoclimatology, Palaeoecology, Quaternary International, Aeolian Research, Marine Geology, PLOS ONE, PEERJ, iScience, Biology, Quaternary, Earth, Journal of Marine Science and Engineering, Heritage, Quaternary Sciences (in Chinese), and Journal of Earth Sciences and Environment (in Chinese)
7. Invited Expert for Mollusk Identification for the Ontario Geological Survey (Canada), and for 14 universities or research institutions in China including Peking University, Beijing Normal University, Lanzhou University, Yunnan University, Henan University, and the Chinese Academy of Sciences
8. Organizing Committee Member of the 2022 INQUA LoessFest
9. Keith Runcorn Travel Award, European Geosciences Union (EGU)
10. Special Funding of the China Postdoctoral Science Foundation
11. CAS President's Award for Excellence (Chinese Academy of Sciences)
12. Third Prize in the Graduate Elite Cup Academic Paper Competition, Jilin University

Projects as Principal Investigator

1. General Program of the National Natural Science Foundation of China (NSFC), Grant No. 42172210, "Study on abrupt climate events during the breeding season recorded by fossil land-snail eggs in the Chinese Loess Plateau since 900 ka", 2022/01–2025/12.
2. General Program of NSFC, Grant No. 41772186, "Climate change during biological breeding seasons recorded by land snails and their fossil eggs over the last four glacial cycles in the Chinese Loess Plateau", 2018/01–2021/12.
3. General Program of NSFC, Grant No. 41372186, "Study on mollusk fossil assemblages of the paleoenvironmental background of early human occupation in the Nihewan Basin from 1.8 to 0.8 Ma", 2014/01–2017/12.
4. General Program of NSFC, Grant No. 41072130, "Ecological and environmental evolution recorded by Middle Miocene land snail fossils in the Chinese Loess Plateau", 2011/01–2013/12.
5. Key Direction Project of the Chinese Academy of Sciences (CAS), Grant No. KZCX2-EW-QN107, "Land snail fossil record of ecological and environmental evolution around 14 Ma in the Chinese Loess Plateau", 2011/01–2013/12.
6. Sub-project of the Strategic Priority Research Program of CAS, Grant No. XDA05130604, "Integrated study on the origin of agriculture", 2011/01–2015/12.
7. Special Project of the Major Research Program on Global Change, Grant No. 2010CB95020402, "Seasonal characteristics of climate during warming processes and quantitative-semi-quantitative reconstruction", 2010/06–2014/12.
8. Special Fund for CAS President's Award for Excellence, "Study on the relationship between Pliocene evolution of land snails and insolation gradient in the Chinese Loess Plateau", 2010/01–2012/12.
9. NSFC Young Scientist Fund, Grant No. 40702030, "Major ecological and environmental evolution recorded by land snail fossils around 8 Ma in the western Chinese Loess Plateau", 2008/01–2010/12.
10. Special Funding of the China Postdoctoral Science Foundation, Grant No. 200801110, "Study on the relationship between East Asian monsoon evolution and solar radiation gradient recorded by Late Miocene snail fossils in the Chinese Loess Plateau", 2008/07–2009/01.
11. First-Class Funding of the China Postdoctoral Science Foundation, Grant No. 20070420066, "Monsoon environmental evolution history recorded by snail fossils from 10 to 6 Ma in the western Chinese Loess Plateau", 2007/09–2008/08.

Projects Participated (Selected)

1. Basic Science Center Project of the National Natural Science Foundation of China (NSFC), "Continental Evolution and Monsoon System Evolution", Grant Nos. 41888101 and 42488201, 2019/01–2028/12.
2. Sub-project of the Strategic Priority Research Program of the Chinese Academy of Sciences (CAS), "Fine processes and quantitative reconstruction of Neogene vegetation succession and events", Grant No. XDB26000000, 2018/06–2022/12.
3. Key Project of NSFC, "Eco-environmental changes and their mechanisms in the Chinese Loess Plateau during low‑eccentricity interglacials of the Quaternary Earth orbital cycles ", Grant No. 41430103, 2015/01–2019/12.
4. Key Project of NSFC, "Microfossil study on the relationship between agricultural origin and environmental change in the Yellow River Basin", Grant No. 41230104, 2013/01–2017/12.
5. Key Direction Project of the CAS Knowledge Innovation Program, "High‑ and low‑latitude interactions recorded by aeolian deposits on different timescales", Grant No. KZCX2-YW-117, 2007/01–2009/12.
6. Key Project of NSFC, "Periodicity of Neogene climate change recorded by aeolian deposits in northern China", Grant No. 40730104, 2008/01–2011/12.
7. General Program of NSFC, "Ecological environment during the Mid‑Pleistocene Transition on the Chinese Loess Plateau and its significance", Grant No. 40672116, 2007/01–2009/12.
8. General Program of NSFC, "Ecological environment of Marine Isotope Stage 11 on the Chinese Loess Plateau and its significance", Grant No. 40372083, 2004/01–2006/12.

1 Wang Yiya, Li Fengjiang, Peng Siyi, Zhao Jiujiang, Zhao Linghao, Dong Yajie, Sun Dongyang, Wu Naiqin, 2026. Feasibility of elemental and microstructural differentiation of land snail eggs from Bradybaena ravida and Cathaica fasciola. Biology, 15(9), 721. https://doi.org/10.3390/biology15090721.

2 Li Fengjiang, Wu Naiqin, Rousseau Denis-Didier, Huang Linpei, Wu Bin, Hao Qingzhen, Dong Yajie, Chen Xiaoyun, 2025. Land-snail eggs from Chinese L9 loess strata and seasonal cooling events during Marine Isotope Stages24–22. Quaternary Science Reviews, 369: 109584.

3 Li Fengjiang, Wu Naiqin, Dong Yajie, Yang Yiquan, Zhang Yueting, Zhang Dan, Hao Qingzhen, Lu Houyuan, 2023. Seasonal climatic instability in the western Chinese Loess Plateau during Marine Isotope Stages 12–10. Scientific Reports, 13: 5725, https://doi.org/10.1038/s41598-023-32923-8.

4 Li Fengjiang, Wu Naiqin, Zhang Dan, Rousseau Denis-Didier, Yang Yiquan, Hao Qingzhen, Dong Yajie, Lu Houyuan, 2022. Glacial-interglacial evolution of seasonal cooling events documented by land-snail eggs from Chinese Loess. Quaternary Science Reviews, 284: 107506.

5 Li Fengjiang, Wu Naiqin, Dong Yajie, Zhang Dan, Zhang Yueting, Huang Linpei, Yang Yiquan, Xu Deke, Zhang Jianping, Lu Houyuan, 2021. Land-snail eggs as a proxy of abrupt climatic cooling events during the reproductive season. Science Bulletin, 66(13): 1274–1277.

6 Li Fengjiang, Wu Naiqin, Rousseau Denis-Didier, Dong Yajie, Zhang Dan, Pei Yunpeng, 2014. Late Miocene-Pliocene paleoclimatic evolution documented by terrestrial mollusk populations in the western Chinese Loess Plateau. PLOS ONE, 9 (4), e95754, 1–12. doi: 10.1371/journal.pone.0095754.

7 Li Fengjiang, Wu Naiqin, Lu Houyuan, Zhang Jianping, Wang Weilin, Ma Mingzhi, Zhang Xiaohu, Yang Xiaoyan, 2013. Mid-Neolithic exploitation of mollusks in the Guanzhong Basin of Northwestern China: Preliminary results. PLOS ONE, 8 (3), e58999, 1–9. doi: 10.1371/journal.pone.0058999.

8 Li Fengjiang, Wu Naiqin, 2010. Pliocene land snail record from western Chinese Loess Plateau and implications for impacts of summer insolation gradient between middle and low latitudes on the East Asian summer monsoon. Global and Planetary Change, 72 (1-2): 73–78.

9 Li Fengjiang, Rousseau Denis-Didier, Wu Naiqin, Hao Qingzhen, Pei Yunpeng, 2008. Late Neogene evolution of the East Asian monsoon revealed by terrestrial mollusk record in western Chinese Loess Plateau: from winter to summer dominated sub-regime. Earth and Planetary Science Letters, 274 (3-4): 439–447.

10 Li Fengjiang, Wu Naiqin, Pei Yunpeng, Hao Qingzhen, Rousseau Denis-Didier, 2006. Wind-blown origin of Dongwan late Miocene-Pliocene dust sequence documented by land snail record in western Chinese Loess Plateau. Geology, 34 (5): 405–408.

11 Li Fengjiang, Wu Naiqin, Denis-Didier Rousseau, 2006. Preliminary study of mollusk fossils in the Qinan Miocene loess-soil sequence in western Chinese Loess Plateau. Science in China, Series D, 49(7): 724–730.

12Wei Fukang, Dong Yajie, Wu Naiqin,Li Fengjiang, Lu Houyuan, 2025.Assessing the temporal stability of snail-climate relationships since the Last Glacial Maximum: Insights from boosted regression trees and LOESS models. Palaeogeography, Palaeoclimatology, Palaeoecology, 662: 112771.

13 Dong Yajie, Wu Naiqin, Li Fengjiang, Lu Houyuan, 2024a. Geographical distributions and ecological ranges of the dominant snail species in China and their palaeoclimatic implications. Palaeogeography, Palaeoclimatology, Palaeoecology, 640: 112108.

14 Dong Yajie, Wu Naiqin, Li Fengjiang, Wei Fukang, Lu Houyuan, 2024b. Compositional changes in land snail assemblages from Chinese loess sequences over the past 20,000 years reveal the unprecedented scale of recent anthropogenic impacts. Palaeogeography, Palaeoclimatology, Palaeoecology, 651: 112388.

15 Dong Yajie, Wu Naiqin, Li Fengjiang, Wei Fukang, Lu Houyuan, 2024. Carbon storage in mollusk shells: An overlooked yet significant carbon sink in terrestrial ecosystems. Science of the Total Environment, 915: 170050.

16 Hao Qingzhen, Peng Shuzhen, Gao Xinbo, Markovic Slobodan B, Li Sheng-Hua, Zhang Junjie, Li Fengjiang, Han Long, Fu Yu, Wu Xuechao, Wang Luo, Xu Bing, Qiao Yansong, Yu Jimin, Guo Zhengtang, 2024. Unusual weakening trend of the East Asian winter monsoon during MIS 8 revealed by Chinese loess deposits and its implications for ice age dynamics. Global and Planetary Change 234: 104389.

17 Xu Deke, Chu Guoqiang, Shen Caiming, Sun Qing, Wu Jing, Li Fengjiang, Dong Yajie, Cui Anning, Wu Naiqin, Lu Houyuan, 2023. 500-Year Periodic Vegetation and Monsoonal Climate Oscillations During the Last Deglaciation in East Asia. Geophysical Research Letters, 50, e2023GL103535.

18 Xu Deke, Lu Houyuan, Chu Guoqiang, Shen Caiming, Sun Qing, Wu Jing, Li Fengjiang, Song Bing, Cui Anning, Li Hao, Wu Naiqin, 2023. Fast response of vegetation in East Asia to abrupt climatic events during the last deglaciation. PNAS Nexus, 2: 1–10

19 Dong Yajie, Wu Naiqin, Li Fengjiang, Zhang Dan, Zhang Yueting, Shen Caiming, Lu Houyuan, 2022. The Holocene temperature conundrum answered by mollusk records from East Asia. Nature Communications, 13: 5153.

20 Dong Yajie, Wu Naiqin, Li Fengjiang, Huang Linpei, Lu Houyuan, Stenseth N.C., 2021. Paleorecords reveal the increased temporal instability of species diversity under biodiversity loss. Quaternary Science Reviews, 269: 107147.

21 Dong Yajie, Wu Naiqin, Li Fengjiang, Lu Houyuan, 2021. Rapid northwestward extension of the East Asian summer monsoon since the last deglaciation: evidence from the mollusk record. Frontiers in Earth Science, 9: 788738.

22 Dong Yajie, Wu Naiqin, Li Fengjiang, Lu Houyuan, 2021. Coupled morphologic and demographic responses of Opeas striatissimum (Gastropoda: Subulinidae) to last deglacial warming. Palaeogeography, Palaeoclimatology, Palaeoecology, 562: 110101.

23 Zhang Dan, Wu Naiqin, Li Fengjiang, Rousseau Denis-Didier, Chen Xiaoyun, Dong Yajie, Lu Houyuan, 2021. Climatic structures and intensities of the last two glacials documented by terrestrial molluscs from Chinese loess sequences. Boreas, 50: 308–320.

24 Dong Yajie, Wu Naiqin, Li Fengjiang, Zhang Dan, Zhang Yueting, Huang Linpei, Chen Xiaoyun, Wu Bin, Lu Houyuan, 2020. Anthropogenic modification of soil communities in northern China for at least two millennia: Evidence from a quantitative mollusk approach. Quaternary Science Reviews, 248, 106579.

25 Dong Yajie, Wu Naiqin, Jiang Wenying, Li Fengjiang, Lu Houyuan, 2020. Cascading response of flora and terrestrial mollusks to last deglacial warming. Global Ecology and Conservation, 24: e01360.

26 Xu Deke, Lu Houyuan, Chu Guoqiang, Shen Caiming, Li Fengjiang, Wu Jing, Wang Luo, Li Hao, Yu Yinghao, Jin Yingyu, Wu Naiqin, 2020. Asynchronous 500-year summer monsoon rainfall cycles between Northeast and Central China during the Holocene. Global and Planetary Change, 195: 103324.

27 Zhang Jianping, Lu Houyuan, Jia Jiwei, Shen Caiming, Wang Shuyun, Chu Guoqiang, Wang Luo, Cui Anning, Liu Jiaqi, Wu Naiqin, Li Fengjiang, 2020. Seasonal drought events in tropical East Asia over the last 60,000 y. PNAS, 117: 30988–30992.

28 Zhang Yueting, Wu Naiqin, Li Fengjiang, Hao Qingzhen, Dong Yajie, Zhang Dan, Lu Houyuan, 2020. Eco-environmental changes in the Chinese Loess Plateau during low-eccentricity interglacial Marine Isotope Stage 19. Science in China Earth Sciences, 63: 1408–1421.

29 Dong Yajie, Wu Naiqin, Li Fengjiang, Chen Xiaoyun, Zhang Dan, Zhang Yueting, Huang Linpei, Wu Bin, Lu Houyuan, 2019. Influence of monsoonal water-energy dynamics on terrestrial mollusk species-diversity gradients in northern China. Science of the Total Environment, 676: 206–214.

30 Xu Deke, Lu Houyuan, Chu Guoqiang, Liu Li, Shen Caiming, Li Fengjiang, Wang Can, Wu Naiqin, 2019. Synchronous 500-year oscillations of monsoon climate and human activity in Northeast Asia. Nature Communications, 10, 4105.

31 Gao Xinbo, Hao Qingzhen, Oldfield Frank, Bloemendal Jan, Deng Chenglong, Wang Luo, Song Yang, Ge Junyi, Wu Haibin, Xu Bing, Li Fengjiang, Han Long, Fu Yu, Guo Zhengtang, 2019. New high-temperature dependence of magnetic susceptibilit-based climofunction for quantifying paleoprecipitation from Chinese loess. Geochemistry, Geophysics, Geosystems, 20: 4273–4291.

32 Wang Can, Lu Houyuan, Gu Wanfa, Wu Naiqin, Zhang Jianping, Zuo Xinxin, Li Fengjiang, Wang Daojing, Dong Yajie, Wang Songzhi, Liu Yanfeng, Bao Yingjian, Hu Yayi, 2019. The development of Yangshao agriculture and its interaction with social dynamics in the middle Yellow River region, China. The Holocene, doi.org/10.1177/0959683618804640.

33 Gao Xinbo, Hao Qingzhen, Wang Luo, Oldfield Frank, Bloemendal Jan, Deng Chenglong, Song Yang, Ge Junyi, Wu Haibin, Xu Bing, Li Fengjiang, Han Long, Fu Yu, Guo Zhengtang, 2018. The different climatic response of pedogenic hematite and ferrimagnetic minerals: Evidence from particle-sized modern soils over the Chinese Loess Plateau. Quaternary Science Reviews, 179: 69–86.

34 Wu Naiqin, Li Fengjiang, Rousseau Denis-Didier, 2018. Terrestrial mollusk records from Chinese loess sequences and changes in the East Asian monsoonal environment. Journal of Asian Earth Sciences, 155: 35–48.

35 Wang Can, Lu Houyuan, Gu Wanfa, Wu Naiqin, Zhang Jianping, Zuo Xinxin, Li Fengjiang, Wang Daojing, Dong Yajie, Wang Songzhi, Liu Yanfeng, Bao Yingjian, Hu Yayi, 2017. The spatial pattern of farming and factors influencing it during the Peiligang culture period in the middle Yellow River valley, China. Science Bulletin, 62: 1565–1568.

36 Dong Yajie, Wu Naiqin, Li Fengjiang, Huang Linpei, Wen Wenwen, 2015. Time-transgressive nature of the magnetic susceptibility record across the Chinese Loess Plateau at the Pleistocene/Holocene transition. PLOS ONE 10(7): e0133541.

37 Xiao Guoqiao, Guo Zhengtang, Dupont-Nivet Guillaume, Lu Houyuan, Wu Naiqin, Ge Junyi, Hao Qingzhen, Peng Shuzhen, Li Fengjiang, Abels Hemmo A, Zhang Kexin, 2012. Evidence for northeastern Tibetan Plateau uplift between 25 and 20 Ma in the sedimentary archive of the Xining Basin, Northwestern China. Earth and Planetary Science Letters, 317-318: 185–195.

38 Zhang Jianping, Lu Houyuan, Wu Naiqin, Li Fengjiang, Yang Xiaoyan, Wang Weilin, Ma Mingzhi, Zhang Xiaohu, 2010. Phytolith evidence for rice cultivation and spread in Mid-Late Neolithic archaeological sites in North-central China. Boreas, 39(3): 592–602.

39 Rousseau Denis-Didier, Wu Naiqin, Pei Yunpeng, Li Fengjiang, 2009. Three exceptionally strong East-Asian summer monsoon events during glacial times in the past 470 kyr. Climate of the Past, 5: 157–169.

40 Wu Naiqin, Chen Xiaoyun, Rousseau Denis-Didier, Li Fengjiang, Pei Yunpeng, Wu Bin, 2007. Climatic conditions recorded by terrestrial mollusc assemblages in the Chinese Loess Plateau during marine Oxygen Isotope Stages 12-10. Quaternary Science Reviews, 26(13-14): 1884–1896.

41 Wu Naiqin, Pei Yunpeng, Lu Houyuan, Guo Zhengtang, Li Fengjiang, Liu Tungsheng, 2006. Marked ecological shifts during 6.2–2.4 Ma revealed by a terrestrial molluscan record from the Chinese Red Clay Formation and implication for palaeoclimatic evolution. Palaeogeography, Palaeoclimatology, Palaeoecology, 233(3-4): 287–299.

42 Pei Yunpeng, Wu Naiqin, Li Fengjiang, 2004. Terrestrial mollusk evidence for the origin and sedimentary environment of the Late Tertiary Red Clay Formation in the Loess Plateau, China. Chinese Science Bulletin, 49(10): 1072–1076.

43李丰江, 杨意权, 吴乃琴, 黄林培, 董亚杰, 2019. 蜗牛卵化石在黄土高原的发现及其对季节内突变气候事件的指示. 第四纪研究, 39(4): 1068–1070.

44 李丰江, 吴乃琴, 董亚杰, 吕厚远, 陈晓云, 张丹, 张月婷, 黄林培, 伍斌, 2016. 黄土高原及周边地区间齿螺（Metodontia）种类的数量分布及其温度和降水量最适范围的定量估算. 第四纪研究, 36(3): 564–574.

45 李丰江, 吴乃琴, Denis-Didier Rousseau, 2006. 黄土高原秦安中新世黄土-古土壤序列的蜗牛化石初步研究. 中国科学(D辑), 36(5): 438–444.

46 李丰江, 吴乃琴, 裴云鹏, 2005. 黄土高原西部秦安新近纪风尘堆积的蜗牛化石证据. 第四纪研究, 25(4): 510–515.

47 李丰江, 邓金宪, 温泉波, 陈广善, 李军敏, 2004. 吉林双辽地区风沙堆积古温度研究. 地理科学, 24(5): 616–619.

48 李丰江, 邓金宪, 温泉波, 陈广善, 2002. 长春腰分水岭黄土剖面常量化学元素特征及其气候意义. 世界地质, 21(1): 53–56.

49 王祎亚，李丰江，赵九江，马生凤，赵令浩，许智超，张保科，孙红宾，孙冬阳，芦苒. 灰巴蜗牛（Bradybaena ravida）卵的特征及温度对卵孵化的影响. 北京农学院学报, 2026, 41(1): 15–20.

50 杨意权, 李丰江, 吴乃琴, 董亚杰, 张丹, 张月婷, 2020. 生物卵化石研究进展简述. 地球环境学报, 11(1): 1–13.

51 吴乃琴, 李丰江, 2008. 陆生蜗牛化石与中国黄土古环境研究. 第四纪研究, 28(5): 831–840.

52 裴云鹏, 吴乃琴, 李丰江, 2004. 晚第三纪红粘土成因和沉积环境的生物学证据——蜗牛化石记录. 科学通报, 49(13): 1294–1298.

53 张月婷, 吴乃琴, 李丰江, 郝青振, 董亚杰, 张丹, 吕厚远, 2020. 低偏心率间冰期(MIS 19)黄土高原生态环境变化及影响机制. 中国科学: 地球科学, 50: 1477–1491.

54 张健平, 吕厚远, 吴乃琴, 李丰江, 杨晓燕, 王炜林, 马明志, 张小虎, 2010. 关中盆地6000～2100 cal. aB.P.期间黍、粟农业的植硅体证据. 第四纪研究, 30(2): 287–297.

55 王灿, 吕厚远, 顾万发, 吴乃琴, 张健平, 左昕昕, 李丰江, 汪道京, 董亚杰, 汪松枝, 刘彦锋, 鲍颖建, 胡亚毅, 2019. 全新世中期郑州地区古代农业的时空演变及其影响因素. 第四纪研究, 39(1): 108–122.